- PhD, University of Reading, 1997. I researched the evolution and development of early vertebrates, focusing on the homeobox genes of the lamprey and amphioxus, in the lab of Peter Holland. Thesis title: Gene duplication in vertebrate evolution (available from Ethos at the British Library).
- BA, Cambridge University, class 2i, 1992. Natural sciences, specialising in zoology (developmental biology, cell biology and animal biology).