Academic qualifications

• PhD, University of Reading, on the evolution and development of early vertebrates, in the lab of Peter Holland (thesis online)
• BA, Cambridge University, natural sciences

Editorial training

• SfEP training courses including on-screen editing and editing medical texts
• Publishing Training Centre courses including advanced copyediting, rewriting and writing and editing for the web
  

Professional memberships

• Advanced Member of the Society for Editors and Proofreaders (SfEP). Membership requires proof of a high degree of competence and substantial experience as an editor, including evidence of training and experience and references from clients.
• Member of the European Association of Science Editors (EASE).